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影片讲述松鼠奎特为了追松果，偶然引发了宇宙事件，改变并威胁着冰川时代的世界。为了拯救自己，话唠树懒希德、猛犸象曼尼、剑齿虎迪亚哥，以及别的动物群族必须离开家园，踏上了他们充满喜剧色彩的冒险旅程，他们来到了充满异国情调的新大陆，并遇到了许多形形色色的新角色。。After suffering a brutal attack on Christmas eve, a young woman, Nikki, struggles to convince her friends that the assailant was none other than Ol' Kris Kringle. When Santa returns to terrorize the group in their remote cabin the next Christmas, Nikki and her friends must overcome disbelief as they fight to stay alive.。夏岚父被韩城权贵陷害入狱，三年前求青梅竹马沈寒之相助遭其母羞辱，沈寒之亦消失。今父病危，夏岚设计接近宋家独子宋望，欲借其权势翻案。沈寒之携误会回国，见她亲近宋望，强势介入，夏岚表面抗拒，内心因旧情动摇。。